Semaeostomeae

Semaeostomeae
Sea nettles, Chrysaora fuscescens
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Cnidaria
Class:	Scyphozoa
Subclass:	Discomedusae
Order:	Semaeostomeae L. Agassiz, 1862
Families
See text

Semaeostomeae (literally "flag mouths") is an order of large jellyfish characterized by four long, frilly oral arms flanking their quadrate mouths. The umbrella is domed with scalloped margins, and the gastrovascular system consists of four unbranched pouches radiating outwards from the central stomach; no ring canal is present. They usually possess eight tentacles; four are per-radical and four are inter-radical.

Taxonomy

The order consists of five families.

• Family Cyaneidae
• Family Drymonematidae
• Family Pelagiidae
• Family Phacellophoridae
• Family Ulmaridae

Differentiation

The three traditional families, Pelagiidae, Cyaneidae, and Ulmaridae, are distinguishable by these characteristics:

• Gastrovascular cavity divided by radial septa into rhopalar and tentacular pouches
  • Pouches simple and unbranched – Pelagiidae
  • Pouches branched – Cyaneidae
• Gastrovascular system in form of unbranched and branching canals, or with anastomosing radial canals – Ulmaridae

In addition, members of the Pelagiidae have no ring canal, and the marginal tentacles arise from umbrella margin. Three genera are in this family.

Gallery

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  "Discomedusae" plate from Ernst Haeckel's Kunstformen der Natur, 1904

References

• Collins, A. G.; Jarms, G.; Morandini, A. C. (2022). "Semaeostomeae". World Register of Marine Species. Retrieved 1 November 2022.
• Ruppert, Edward E.; and Fox, Richard S. (1988). Seashore Animals of the Southeast, University of South Carolina Press, 273. ISBN 0-87249-535-3.
• Hayward, Peter J.; and Ryland, John S. (1995). Handbook of the Marine Fauna of North-West Europe, Oxford University Press, 65. ISBN 0-19-854055-8.
• Bayha K., Dawson M. (2010) New family of allomorphic jellyfishes, Drymonematidae (Scyphozoa, Discomedusae), emphasizes evolution in the functional morphology and trophic ecology of gelatinous zooplankton. Biological Bulletin 219: 249–267.